Is it okay to use PCI Gen 4 drives with gen 3 slots?

Apakah slot SSD gen 3 bisa dipasangkan dengan SSD gen 4? can i use gen 4 ssd on gen 3 slot
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can you apply for dv lottery while in usa 2_1 slot on my ASUS ROG Strix x570i mobo The only thing I can find in bios is set to _1 link = Gen 4 Yet the SSD is still running at Gen 3